Интенсивность колебаний

Это приспособление позволяло предложить скорпиону следующую дилемму. В естественной ситуации ноги скорпиона, которые находятся ближе к источнику колебаний, стимулируются раньше и сильнее. На разделенной пополам площадке можно было дифференцировать время и интенсивность колебаний: например, подавать к правым ногам импульс раньше, но вдвое слабее, чем к левым ногам. Скорпион должен был выбрать то, что для него важнее. Эксперимент повторяли многократно, и скорпион неизменно поворачивался направо независимо от интенсивности стимула, поступавшего к левым ногам. Следовательно, для установления направления на источник колебаний скорпион руководствуется именно временем получения сигнала, а не его относительной интенсивностью. Следует отметить, что скорпион безошибочно определял, какие ноги получали сигнал первыми. Судя по быстроте и точности поворотов скорпиона вправо или влево, оптимальное время запаздывания сигнала составляет 1—2 мс. За одну миллисекунду поверхностная волна проходит 5 см, т.е. как раз пересекает сенсорное поле скорпиона. Однако реакции скорпиона остаются достаточно четкими даже при запаздывании всего на 0,2 мс. Остается выяснить, как скорпион определяет расстояние до жертвы. Как показали наши наблюдения в природе, скорпион чувствует добычу уже на расстоянии около 15 см, а если до нее не более 10 см, то он ошибается крайне редко. К сожалению, из-за того, что скорпионы отказываются охотиться в лабораторных условиях, не удается исследовать механизм оценки расстояния до жертвы, так что в этом вопросе мы пока находимся в области предположений. Можно лишь сказать, что при определении расстояния, так же как и при определении направления, должны восприниматься различия во времени поступления и в амплитуде сигналов, приходящих к разным рецепторам. Вполне вероятно также, что та информация, которую несут волны сжатия и которая, по-видимому, не нужна для реакций ориентации, используется скорпионом для определения расстояния до добычи. Можно предположить, что скорпион регистрирует разницу во времени стимуляции тарзальных волосков волнами сжатия и шелевидной сенсиллы медленнее распространяющимися рэлеевскими волнами, а эта величина должна быть пропорциональна расстоянию до жертвы. Возможен и другой механизм, основанный на восприятии затухания сигналов.

 

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